1. Chemistry Research Laboratory and the Oxford Centre for Integrative Systems Biology, University of Oxford Mansfield Road Oxford OX1 3TA UK
2. Henry Wellcome Building for Molecular Physiology, University of Oxford, Roosevelt Drive Oxford OX3 7BN UK
3. Institut für Tierökologie und Zellbiologie, Tierärztliche Hochschule Hannover, Bünteweg 17d Hannover 30559 Germany
4. Department of Zoology, University of Oxford, South Parks Road Oxford OX1 3PS UK