1. Department of Soil Science, The University of Reading, Whiteknights, PO Box 233, Reading, RG6 6DW;
2. Soil‐Plant Dynamics Unit, Scottish Crop Research Institute, Dundee, DD2 5DA;
3. SIMBIOS Centre, School of Science and Engineering, University of Abertay Dundee, Bell St., Dundee DD1 1HG;
4. Department of Plant and Soil Science, University of Aberdeen, Aberdeen AB24 3UU