1. Institute of Rural Studies, Llanbadarn Fawr , University of Wales, Aberystwyth, Ceredigion SY23 3AL, UK;
2. Conservation and Ecology Department, SAC Aberdeen, Craibstone, Bucksburn, Aberdeen AB2 9TS, UK;
3. present address: University of Brighton, Department of Pharmacy, Cockcroft Building, Lewes Road, Brighton, BN2 4GJ, UK