1. Both authors contributed equally to this work.
2. Laboratoire de Génétique Moléculaire de la Neurotransmission et des Processus Neurodégénératifs, CNRS‐UMR C9923, Hôpital de la Pitié Salpêtrière, Paris, France,
3. Center for Genome Research, AFRC, University of Edinburgh, Edinburgh, UK,
4. Present address: National Institute of Medical Research, the Ridgeway, Mill Hill, London, UK,
5. Laboratoire de Neurobiologie Cellulaire et Moléculaire, CNRS, Gif‐sur‐Yvette, France,
6. Present address: Laboratoire de Pharmacogénétique et Abords Thérapeutiques des Maladies Héréditaires, INSERM U458, Hôpital Robert Debré, Paris, France,
7. Present address: Department of Anatomy and Developmental Biology, Royal Free Hospital School of Medicine,Rowland Hill Street, London, UK