1. Department of Plant Pathology and Microbiology, Horticulture Research International, Wellesbourne, Warwick, CV35 9EF, UK;
2. Department of Soil and Environment Sciences, Horticulture Research International, Wellesbourne, Warwick, CV35 9EF, UK;
3. Department of Animal and Plant Sciences, University of Sheffield, Sheffield, S10 2TN, UK