1. Dauphin Island Sea Laboratory, 101 Bienville Blvd, Dauphin Island, AL 36528 USA;
2. University of South Alabama, Department of Marine Sciences, Mobile AL USA;
3. University of Alabama, Department of Biological Sciences, Tuscaloosa, AL USA;
4. U. S. Environmental Protection Agency, Gulf Ecology Division, Gulf Breeze, FL USA 32561