1. School of Biological Science, University of New South Wales, NSW 2052, Australia
2. Queensland Parks and Wildlife Service, P.O. Box 155, Albert Street, QLD 4002 Australia
3. Biodiversity Survey and Research Division, New South Wales National Parks and Wildlife Service, P.O. Box 1967, Hurstville, NSW 2220, Australia
4. Faculty of Arts, Health and Sciences, Central Queensland University, Rockhampton, QLD 4702, Australia
5. Department of Conservation Biology, Daniel F. and Ada L. Rice Center, Chicago Zoological Society,
Brookfield, IL 60513, U.S.A.