1. School of Natural Resources and Environment, The University of Michigan, Ann Arbor, MI 48109–1115, U.S.A.
2. Pacific Meridian, 455 East Eisenhower Parkway, Ann Arbor, MI 48108, U.S.A.
3. Department of Ecology and Evolutionary Biology, Cornell University, Ithaca, NY 14853, U.S.A.
4. Departamento de Biología, Universidad de Los Andes, Mérida 5251, Venezuela
5. Museo de Zoologia, Universidad Nacional Experimental de los Llanos Occidentales Ezequiel Zamora,
Guanare, Estado Portuguesa 3310, Venezuela