1. Department of Life Science, Pohang University of Science and Technology, Hyojadong San 31, Pohang 790–784, Republic of Korea
2. School of Environmental Engineering, Pohang University of Science and Technology, Hyojadong San 31, Pohang 790–784, Republic of Korea
3. Department of Molecular and Cell Biology, The University of Connecticut, Storrs, CT 06269, USA
4. Plant Molecular Biology and Biotechnology Research Center, Chinju 660–701, Republic of Korea