1. 1 Laboratory of Plant Molecular Genetics, Nara Institute of Science and Technology, 8916–5 Takayama, Ikoma 630–0101, Japan, 2 Laboratory of Bioorganic and Agricultural Chemistry, School of Agriculture, Ibaraki University, Ami 300–0332, Japan, and 3 Faculty of Agriculture, Kyushu University, Hakozaki, Fukuoka 812–0053, Japan