1. Animal Science and Microbiology Department, Institute of Grassland and Environmental Research, Plas Gogerddan, Aberystwyth, Ceredigion SY23 3EB, UK;
2. Agrotechnological Research Institute, Postbus 17, 6700 AA Wageningen, The Netherlands and
3. Biochemistry and Physiology Department, IACR‐Rothamsted, Harpenden, Herts AL5 2JQ, UK