1. GIROQ, Département de biologie, Université Laval, Ste‐Foy, Quebec G1K 7P4, Canada,
2. Virginia Cooperative Fish and Wildlife Research Unit, Virginia Polytechnic Institute and State University, Blacksburg, Virginia 24061–0321, USA,
3. Department of Biology, McGill University, Montréal, Quebec H3 A 1B1, Canada