1. Centre for Plant Diversity and Systematics, Department of Botany, School of Plant Sciences, The University of Reading, Whiteknights, Reading RG6 6AS,,
2. Conservation Projects Development Unit, Living Collections Department, The Royal Botanic Gardens, Kew, Richmond, Surrey TW9 3AB, UK,,
3. CONAF Region del Bio Bio, Barros Arana 215, Concepción, Chile,,
4. Department of Agricultural Botany, School of Plant Sciences, The University of Reading, Whiteknights, Reading RG6 6AS, UK