1. Department of Ecology, Evolution, and Behavior, University of Minnesota, 100 Ecology Bldg, 1987 Upper Buford Circle, St. Paul, MN 55108,
2. Ecology, Evolution, and Marine Biology, University of California, Santa Barbara, CA 93106,
3. Geochemistry Department, University of California, Irvine, CA 92717‐3100,
4. Center for Atmospheric Sciences, University of California, Berkeley, CA 94702, Institute for Arctic Biology, University of Alaska, Fairbanks, AK 99775, USA