1. School of Botany and Zoology, Australian National University, Canberra, ACT 0200, Australia,
2. Centre for Identification and Diagnostics, School of life Sciences, The University of Queensland, Brisbane, QLD 4072, Australia,
3. Conservation and Horticulture, Royal Botanic Gardens Sydney, Mount Annan Botanic Garden, Mount Annan, NSW 2567, Australia