1. Aquatic Ecology Group, Division of Ecology, Department of Biological Sciences, University of Calgary, Calgary, Alberta, Canada T2N 1N4,
2. National Hydrology Research Institute, 11 Innovation Boulevard, Saskatoon, Saskatchewan, Canada S7N 3H5,
3. Sustainable Forest Management Network of Centres of Excellence, G‐208, Biological Sciences Building, University of Alberta, Edmonton, Alberta, Canada T6G 2E9