1. Pelagic Fisheries Research Program, Joint Institute for Marine and Atmospheric Research, University of Hawaii, 2570 Dole Street, Honolulu, Hawaii 96822, USA,
2. Honolulu Laboratory, Southwest Fisheries Science Center, NMFS, NOAA, 2570 Dole Street, Honolulu, Hawaii 96822, USA,
3. Present address: CSIRO, GPO Box 1538, Hobart, TAS 7001, Australia