1. Département de Sciences Biologiques, Université de Montréal, C.P. 6128, Succursale Centre-ville, Pavillon Marie-Victorin, Montreal, Quebec H3C 3J7, Canada, Department of Geosciences, Guyot Hall, Princeton University, Princeton, New Jersey, 08544, and Departments of Chemistry and Atmospheric and Oceanic Sciences, McGill University, 801 Sherbrooke Street West, Montreal, Quebec H3A 2K6, Canada