1. Faculty of Advanced Life Science and Graduate School of Life Science, Hokkaido University,
N21, W11, Kita-ku, 001-0021 Sapporo, Japan
2. Immunology Unit & Research Center for Emerging Infections and Zoonoses (RIZ), University of Veterinary Medicine Hannover, Bünteweg 17, 30559 Hannover, Germany
3. Medicinal Chemistry Pharmaceuticals, Co., Ltd. N9W15,
Chuo-ku, 060-0009 Sapporo, Japan