1. Florey Institute of Neuroscience and Mental Health, University of Melbourne, Melbourne, Victoria 3010, Australia
2. Walter and Eliza Hall Institute of Medical Research, Parkville, Victoria 3052, Australia
3. Department of Surgery, The University of Melbourne, The Royal Melbourne Hospital, University of Melbourne, Parkville, Victoria 3010, Australia
4. Department of Anatomical Pathology, Alfred Hospital, Prahran, Victoria 3004, Australia
5. Department of Biochemistry, Emory University School of Medicine, Atlanta, Georgia 30322, United States
6. Department of Neurology, Emory University School of Medicine, Atlanta, Georgia 30322, United States