1. RIKEN Center for Brain Science, RIKEN, Wako City, Saitama 351-0198, Japan
2. School of Pharmaceutical Sciences (Shenzhen), Sun Yat-sen University, Guangzhou 510275, China
3. Graduate School of Science, Osaka University, Toyonaka, 560-0043, Japan
4. RIKEN Cluster for Pioneering Research, RIKEN, Wako City, Saitama 351-0198, Japan