1. Barts Health NHS Trust, LondonE1 2ES, U.K.
2. Department of Pediatrics, Neurological Sciences, and Anatomy and Cell Biology, Rush University Medical Center, Chicago, Illinois60612, United States
3. Molecular and Human Genetics, Baylor College of Medicine, Houston, Texas77030, United States
4. Department of Molecular Biology, Massachusetts General Hospital, Boston, Massachusetts02114, United States
5. Department of Genetics, Harvard Medical School, Boston, Massachusetts02115, United States
6. Simons Initiative for the Developing Brain, University of Edinburgh, EdinburghEH8 9XD, U.K.