1. School of Natural Resources and Environment, University of Michigan, 440 Church St., Ann Arbor, Michigan 48109, United States
2. Lake Michigan Field Station, Great Lakes Environmental Research Laboratory, NOAA, 1431 Beach St. Muskegon, Michigan 49441, United States
3. Department of Biology, Hope College, Holland, Michigan 49423, United States
4. Graham Environmental Sustainability Institute, University of Michigan, Ann Arbor, Michigan 48109, United States