1. Institute for Protein Research, Osaka University, and CREST, Japan Science and Technology Agency, Yamadaoka 3-2, Suita, Osaka 565-0871, Japan, Institute of Scientific and Industrial Research, Osaka University, Mihogaoka, Ibaraki, Osaka 567-0047, Japan, and Department of Pathological Sciences, Faculty of Medical Sciences, University of Fukui, and CREST, Japan Science and Technology Agency, Matsuoka, Fukui 910-1193, Japan