1. Institute of Biomedical Sciences, Academia Sinica, Taipei 11529, Taiwan
2. Department of Biological Sciences and Technology, National Chiao Tung University, Hsin-Chu 1001, Taiwan
3. Institute of Cellular and Organismic Biology, Academia Sinica, Taipei 11529, Taiwan
4. Graduate Institute of Medicine, College of Medicine, Kaohsiung Medical University, Kaohsiung 80708, Taiwan
5. Department of Biochemistry, Faculty of Medicine, The Hebrew University, Jerusalem 91120, Israel