1. State Key Laboratory of Herbage Improvement and Grassland Agroecosystems, College of Ecology, Lanzhou University, Lanzhou 730000, China
2. Key Laboratory of Microbial Resources Exploitation and Application of Gansu Province, Institute of Biology, Gansu Academy of Sciences, Lanzhou, Gansu 730000, China
3. The UWA Institute of Agriculture, and School of Agriculture and Environment, The University of Western Australia, Perth, WA 6001, Australia
4. Ottawa Research and Development Centre, Agriculture and Agri-Food Canada, Ottawa K1A 0C6, Canada