1. Department of Bioscience Biotechnology, Faculty of Agriculture, Shinshu University, Nagano 399-4598 Japan
2. Mizkan Company, Ltd., 2-6 Nakamura, Handa, Aichi 475-8585, Japan
3. Department of Bioscience and Food Production Science, Interdisciplinary Graduate School of Science and Technology, Shinshu University, Nagano 399-4598, Japan