1. Department of Biology, University of Ottawa, 30 Marie Curie Street, P.O. Box 450, Station A, Ottawa, Ontario K1N 6N5, Canada; Agriculture and Agrifood Canada, Ottawa, Ontario K1A 0C6, Canada; and Département des sciences biologiques, Université du Québec à Montréal, C.P. 8888, Succursale Centre-ville, Montréal, Québec H3C 3P8, Canada