1. National Institute of Fruit Tree Science, Okitsu, Shimizu, Shizuoka, 424-0292, Japan, Genomic Sciences Center, RIKEN, Hirosawa, Wako, Saitama, 351-0198, Japan, Wakayama Prefectural Museum of Natural History, Funao, Kainan, Wakayama, 642-0001, Japan, and Graduate School of Agricultural and Life Sciences, The University of Tokyo, Bunkyo-ku, Tokyo, 113-8657, Japan