1. State Key Laboratory of Pulp and Paper Engineering, South China University of Technology (SCUT), Guangzhou, Guangdong 510640, China
2. Institute for Bioscience and Biotechnology Research (IBBR), University of Maryland, College Park, Maryland 20742, United States
3. School of Resource and Environmental Science, Hubei Biomass-Resource Chemistry and Environmental Biotechnology Key Laboratory, Wuhan University, Wuhan 430079, China