1. Key Laboratory of Forest Plant Ecology, Ministry of Education, Northeast Forestry University, 26 Hexing Road, Xiangfang District, Harbin 150040, P. R. China
2. Engineering Research Center of Forest Bio-preparation, Ministry of Education, Northeast Forestry University, 26 Hexing Road, Xiangfang District, Harbin 150040, P. R. China
3. College of Chemistry, Chemical Engineering and Resource Utilization, Northeast Forestry University, 26 Hexing Road, Xiangfang District, Harbin 150040, P. R. China
4. The College of Forestry, Beijing Forestry University, 35 Qinghua Dong Lu, Haidian District, Beijing 100083, China