1. Laboratory of Biotechnology, Dalian Institute of Chemical Physics, CAS, 457 Zhongshan Road, Dalian 116023, PR China
2. Dalian Key Laboratory of Energy Biotechnology, Dalian Institute of Chemical Physics, CAS, 457 Zhongshan Road, Dalian 116023, PR China
3. Department of Microbiology, Faculty of Science, Kasetsart University, Bangkok 10900, Thailand
4. School of Bioengineering, Dalian University of Technology, 2 Linggong Road, Dalian 116024, China