1. Graduate School of Medical Life Science, Yokohama City University, Yokohama, Kanagawa 230-0045, Japan
2. RIKEN Center for Biosystems Dynamics Research, Yokohama, Kanagawa 230-0045, Japan
3. Faculty of Medicine, Dentistry, and Pharmaceutical Sciences, Okayama University, Okayama 700-8530, Japan
4. Graduate School of Science, Chiba University, Inage, Chiba 263-8522, Japan