1. RIKEN Center for Biosystems Dynamics Research, 6-7-3 Minatojima Minamimachi,
Chuo-ku, Kobe, Hyogo 650-0047, Japan
2. RIKEN Center for Life Science Technologies, 6-7-3 Minatojima Minamimachi, Chuo-ku, Kobe, Hyogo 650-0047, Japan
3. Department of In Vivo Imaging, Tokushima University, 3-18-15 Kuramoto-Cho, Tokushima, Tokushima 770-8503, Japan
4. Human Brain Research Center, Graduate School of Medicine, Kyoto University, 54 Shogoin-Kawahara-Cho, Sakyo-ku, Kyoto 606-8507, Japan