1. RIKEN Plant Science Center, 1-7-22 Suehiro-cho, Tsurumi-ku, Yokohama 235-0045, Graduate School of Nanobiosciencs, Yokohama City University, 1-7-29 Suehirocho, Tsurumi-ku, Yokohama 230-0045, and Graduate School of Bioagricultural Sciences and School of Agricultural Sciences, Nagoya University, 1 Furo-cho, Chikusa-ku, Nagoya-shi 464-8601, Japan