1. Department of Ocean Sciences and Training Vessel Umitaka-maru, Tokyo University of Fisheries, Minato-ku, Tokyo 108-8477, Japan, Environment and Safety Department, Konica Corporation, Sakura-machi, Hino, Tokyo 191-8511, Japan, and Institute of Molecular and Cellular Biosciences, The University of Tokyo, Bunkyo-ku, Tokyo 113-8657, Japan