1. Institute for Biological Sciences, Steacie Institute for Molecular Sciences, and Institute for Microstructural Sciences, National Research Council Canada, Ottawa, Ontario, Canada K1A 0R6, Department of Environmental Biology, Ontario Agricultural College, University of Guelph, Guelph, Ontario, Canada N1G 2W1, Department of Biochemistry, Microbiology and Immunology, University of Ottawa, Ottawa, Ontario, Canada K1N 6N5