1. College of Biotechnology & Food Science, Tianjin University of Commerce, Tianjin 300134, China
2. Tianjin Key Laboratory of Food Biotechnology, Tianjin 300134, China
3. College of Biotechnology, Tianjin University of Science &Technology, Tianjin 300457, China
4. Key Laboratory of Industrial Fermentation Microbiology, Ministry of Education, Tianjin 300457, China