1. CAS Key Laboratory of Coastal Environmental Processes and Ecological Remediation, Shandong Provincial Key Laboratory of Coastal Environmental Processes, YICCAS, Yantai Institute of Coastal Zone Research (YIC), Chinese Academy of Sciences (CAS), Yantai Shandong 264003, P. R. China
2. University of Chinese Academy of Sciences, Beijing 100049, P.R. China
3. Laboratory for Marine Biology and Biotechnology, Qingdao National Laboratory for Marine Science and Technology, Qingdao, Shandong 266237, P.R. China
4. Center for Ocean Mega-Science, Chinese Academy of Sciences, Qingdao, Shandong 266071, P.R. China