1. Department of Biophysics, Graduate School of Science, Kyoto University, Kyoto, Japan
2. Cellular Informatics Laboratory, RIKEN, Wako, Saitama, Japan
3. Laboratory for Cell Signaling Dynamics, RIKEN Quantitative Biology Center, Suita, Osaka, Japan
4. Laboratory of Organic Chemistry for Life Science, Kobe Pharmaceutical University, Kobe, Hyogo, Japan
5. Research Organization for Science and Technology, Ritsumeikan University, Kusatsu, Shiga, Japan