1. The Institute of Physical and Chemical Research (RIKEN), Wako, Saitama 351-0198, Japan, Precursory Research for Embryonic Science and Technology (PRESTO), Japan Science and Technology Corporation (JST), and Institute of Molecular and Cellular Biosciences, The University of Tokyo, Bunkyoku, Tokyo 113-0032, Japan, and Yuki Research Center, Nihon Bayer Agrochem K. K., Yuki, Ibaraki 307-0001, Japan