1. Graduate School of Science, Hokkaido University N21W10, Sapporo 001-0021, Japan, Research Institute for Electronic Science, Hokkaido University N21W10, Sapporo 001-0021, Japan, Department of Molecular Pathobiology, Research Center for Zoonosis Control, Hokkaido University N20W10, Sapporo 001-0020, Japan, 21st Century COE Program for Zoonosis Control, Hokkaido University N20W10, Sapporo 001-0020, Japan, and CREST, JST, 4-1-8 Honcho Kawaguchi, Saitama 332-0012, Japan