1. Institute of Zoology, Chinese Academy of Sciences, Beijing, 100101, People’s Republic of China, Department of Biology and Chemistry, City University Hong Kong, Tat Chee Avenue, Kowloon, Hong Kong SAR, People’s Republic of China, Environmental Measurement Group, National Institute of Advanced Industrial Science and Technology, Onogawa 16-1, Tsukuba, Ibaraki 305-8569, Japan, and Siberia Tiger Park Heilongjiang, Harbin, 88 Songbei Road, 150028, People’s Republic of China