1. Department of Botany, University of British Columbia, #3529-6270 University Boulevard, Vancouver, BC, Canada V6T 1Z4, Agriculture & Agri-Food Canada, Pacific Agri-Food Research Centre, Highway 97, Summerland, BC, Canada V0H 1Z0, and Department of Pathology and Laboratory Medicine, University of British Columbia, 2733 Heather Street, C-Floor, Vancouver, BC, Canada V5Z 1M9