1. Department of Wood Science, The University of British Columbia, 2900-2424 Main Mall, Vancouver, BC, Canada V6T 1Z4, Department of Cell and Systems Biology, University of Toronto, 25 Willcocks Street, Toronto, Ontario, Canada M5S 3B2, Chair of Advanced Papermaking, British Columbia Institute of Technology, 3700 Willingdon Avenue, Burnaby, BC, Canada V5G 3H2