1. Graduate School of Life Sciences, Tohoku University, Aoba-ku, Sendai 980-8577, Japan
2. Frontier Research Institute for Interdisciplinary Sciences, Tohoku University, Aoba-ku, Sendai 980-8577, Japan
3. Cell Biology Center, Institute of Innovative Research, Tokyo Institute of Technology, Yokohama, Kanagawa 226-8503, Japan
4. Laboratory for Chemistry and Life Science, Institute of Innovative Research, Tokyo Institute of Technology, Yokohama, Kanagawa 226-8503, Japan