1. Institute
of Marine Sciences, School of Biological Sciences, University of Portsmouth, Ferry Road, Portsmouth, Hampshire PO4 9LY, United Kingdom
2. Environmental
Research Institute, North Highland College, UHI Millennium Institute, Castle Street, Thurso, Caithness, Scotland KW14 7JD, United Kingdom
3. School
of Biosciences, University of Wales Cardiff, Main Building, Museum Avenue, Cardiff, Wales CF10 3TL, United Kingdom