1. Department of Physics and Biophysics, University of San Diego, 5998 Alcala Park, San Diego, California 92110, United States
2. School of Physics and Astronomy, University of Edinburgh, Peter Guthrie Tait Road, Edinburgh, EH9 3FD, United Kingdom
3. MRC Human Genetics Unit, Institute of Genetics and Molecular Medicine University of Edinburgh, Edinburgh EH4 2XU, United Kingdom