1. National Research Council Canada, 1200 Montreal Road, Ottawa, Canada ON K1A 0R6, University of Ottawa, Eye Institute, 501 Smyth Road, Ottawa, Canada ON K1H 8L6, University of Tennessee Health Center, Memphis, Tennessee, University of Aberdeen, Department of Ophthalmology, Institute of Medical Science, Aberdeen, Scotland AB25 2ZD, and University of Ottawa, Department of Ophthalmology, 451 Smyth Road, Ottawa, Canada K1H 8M5