1. Department of Evolutionary and Functional Biology, University of Parma, Viale G. P. Usberti 11/A, I-43100 Parma, Italy; Department of Evolutionary and Experimental Biology, University of Bologna, Via Irnerio 42, I-40126 Bologna, Italy; and Department of Biology and Evolution, University of Ferrara, C.so Ercole d’Este 32, I-44100 Ferrara, Italy